2.2 Pre-disposition to culture
In the late twentieth century humans developed a new respect for chimpanzee culture. A survey published in 1999 listed no fewer than thirty-nine items of chimpanzee behavior in the wild that seemed to be culturally transmitted. For example, some chimpanzee populations have a technique for cracking nuts with stones-more precisely, they place the nut on one stone and break it with another. Chimpanzees are not genetically programmed to do this; all they get from their genes is the capacity to acquire the skill, normally by learning it from other chimpanzees. This kind of evidence makes it abundantly clear that humans are not the only animals that have culture.
Yet by human standards, the culture of chimpanzees is very limited. They do not seem to develop it to any significant degree. I what we see now is all they have accumulated over an indefinitely long period, then by our standards it is not very much. Humans domesticate the horse and invent the wheel; they put them together and have a horse and cart; eventually they replace the horse with an internal combustion engine and have a car. In the same vein we might imagine nutcracking chimpanzees substituting a third stone for the nut and shaping a stone tool; this is something hominid toolmakers did. But chimpanzees have not done this, and in general it does not seem that one item of their culture serves as a platform for the development of another. In sum, chimpanzee culture is neither cumulative, no dynamic.
The climatic instability of the Pleistocene made it a good time for cultural agility, so also the climatic stability of the Holocene made it a good time for cultural accumulation. It was not that Neolithic cultures were longlasting: typically their life spans are measured in centuries, in contrast to the millennia of Upper Palaeolithic cultures. But the basic innovations were conserved from one culture to another, with no need for anyone to waste time reinventing the wheel. Humans took advantage of the Holocene ecological opportunity because of a genetic predisposition to culture which enable them early in the Holocene to come up with farming.  This was the greatest cultural platform since the emergence of the species.